Geologicheskii Vestnik

The study of the Late Neopleistocene and Holocene molluscs from caves of the Southern Urals demonstrated that, despite the fact that the complexes in different periods are represented by similar species of molluscs, the predominance of some species over others and mollusc shells numbers make it possible to reconstruct the palaeoecological environments of the past. It was discovered that molluscs during Tabulda time (MIS 3) inhabited forest biotopes in a temperate climate and moderate humidity; in some places on the slopes, there were steppe and arid areas; in the Kudashevo time (MIS 2), molluscs lived in a moderately cold climate among shrubs, in woodlands, on open slopes. During the early Holocene, in a temperate climate, molluscs were predominantly found in open spaces — on the rocks of southern slopes and in woodlands with arid or slightly humid conditions; open spaces were widespread in the Middle Holocene; molluscs lived in meadows, valley forests and rocky slopes covered with light forests, the climate was temperate. In the Late Holocene, under conditions of a temperate and humid climate, molluscs inhabited mainly stony slopes with woodlands and open meadows, where conditions were more arid, less often they were found among shrubs and in woodlands under conditions of high humidity. Malacofaunistic data on the reconstruction of the habitat in the Late Pleistocene and Holocene are confirmed by palynological studies.

molluscs, caves, Late Neopleistocene, Holocene, palaeoecology, Southern Urals

• Adam W. Faune de Belgique: Mollusques, 1 / Molluscque Terrestres et Dulcicoles. Institut Royal des Sciences Naturelles de Belgique. 1960. 402 p.
• Alexandrowicz W., Boguckyj A., Dmytruk R., Łanczont M. Malakofauna lessow Naddniestrza Halickiego // Studia geologica Polonica. 2002. V. 119. P. 253–290.
• Alimbekova L.I., Danukalova G.A., Epifanova M.S. Itogi izucheniya sporovo-pyl’tsevykh spektrov po shurfu peshchery “Zapovednaya” // Yezhegodnik — 1996. Informatsionnyye materialy / UNTs RAN. 1998. P. 8–12 (in Russian).
• Animal base, n.d. www.animalbase.uni-goettingen.de. Cohen K.M., Gibbard P.L. Global chronostratigraphical correlation table for the last 2.7 million years, version 2019 QI-500 // Quaternary International. 2019. V. 500. P. 20–31. DOI:10.1016 / j.quaint.2019.03.009
• Danukalova G.A. Utochnennaya regional’naya stratigraficheskaya skhema kvartera Predural’ya i osnovnyye sobytiya na territorii Yuzhno-Ural’skogo regiona // Stratigrafiya. Geologicheskaya korrelyatsiya. 2010. T. 18. №3. P. 1–18 (in Russian).
• Danukalova G.A., Alimbekova L.I. Sporovo-pyl’tsevaya kharakteristika verkhnepleystotsenovykh otlozheniy peshchery “Sikiyaz-Tamak 7” // Yezhegodnik — 1997. Informatsionnyye materialy / UNTs RAN. 1999. P. 53–57 (in Russian).
• Danukalova G., Yakovlev A. A review of biostratigraphical investigations of palaeolithic localities in the Southern Urals region // Quaternary International. 2006. V. 149 (1). 37–43. DOI:10.1016/j.quaint.2005.11.016
• Danukalova G.A., Yakovlev A.G., Alimbekova L.I., Kosintsev P.A., Morozova E.M., Eremeyev A.A. Biostratigrafiya chetvertichnykh otlozheniy peshcher i rechnykh terras shirotnogo techeniya reki Beloy // Ekologicheskiye aspekty Yumaguzinskogo vodokhranilishcha. Ufa: Gilem, 2002. P. 32–57 (in Russian).
• Danukalova G., Yakovlev A., Alimbekova L., Yakovleva T., Morozova E., Eremeev A., Kosintsev P. Biostratigraphy of the Upper Pleistocene (Upper Neopleistocene) — Holocene deposits of the Lemeza River valley of the Southern Urals region (Russia) // Quaternary International. 2008. V. 190 (1). P. 38–57. DOI.org / 10.1016 / j.quaint.2008.04.003
• Danukalova G., Yakovlev A., Osipova E., Yakovleva T., Kosintsev P. Biostratigraphy of the late Upper Pleistocene (upper Neopleistocene) to Holocene deposits of the Belaya River valley (southern Urals, Russia) // Quaternary International. 2011. V. 231 (1–2). P. 28–43. DOI: 10.1016/j.quaint.2010.06.034
• Danukalova G., Lefort J.‑P., Osipova E., Monnier J.‑L. Recent advances in the stratigraphy of the Upper Pleistocene of Westernmost Europe: La Haute Ville and Bréhat Cliffs (Northern Brittany, France) // Quaternary International. 2013. V. 284. P. 30–44. DOI: 10.1016 / j.quaint.2011.10.027
• Danukalova G.A., Osipova E.M., Khenzykhenova F.I. Pozdnepleystotsenovaya fauna mollyuskov (KIS 3) iz mestonakhozhdeniya Bol’shoy Naryn (Baykal’skiy Region, Vostochnaya Sibir’, Rossiya) // Geologicheskiy sbornik, №10. Informatsionnyye materialy / IG UNTs RAN. Ufa: OOO “DizaynPoligrafServis”. 2013. P. 99–108 (in Russian).
• Danukalova G., Osipova E., Khenzykhenova F., Sato T. The molluscs record: A tool for reconstruction of the Late Pleistocene (MIS 3) palaeoenvironment of the Bol’shoj Naryn site area (Fore-Baikal region, Eastern Siberia, Russia) // Quaternary International. 2015. V. 355. P. 24–33. DOI: 10.1016 / j.quaint.2014.08.034
• Danukalova G.A., Yurin V.I., Kosintsev P.A., Osipova E.M., Kurmanov R.G. Biostratigraficheskiye issledovaniya otlozheniy verkhnego pleystotsena i golotsena peshchery Sikiyaz-Tamak 7 (Yuzhnyy Ural, Rossiya) // Geologicheskiy vestnik. 2018. №1. P. 144–161. DOI:10.31084 / 2619–0087 / 2018‑1‑13 (in Russian).
• Danukalova G., Kosintsev P., Yakovlev A., Yakovleva T., Osipova E., Kurmanov R., Kolfschoten T., Izvarin E. Quaternary deposits and biostratigraphy in caves and grottoes located in the Southern Urals (Russia) // Quaternary International. 2020. V. 546. P. 84–124. DOI:10.1016/j.quaint.2020.02.007
• Falkner G., Ripken T.E.J., Falkner M. Mollusques continentaux de France. Liste du references annotees et Bibliographie. Patrimoines naturels 52. Museum National d’ Histoire Naturelle (Paris). France, 2002. 350 p.
• Germain L. Faune de France. Mollusques terrestres et fluviatiles. T. 21 et 22. Paris: Lechevalier, 1930. 893 pp.
• Kerney M.P., Cameron R.A.D. Guide des Escargots et limaces d’Europe. — Lausanne: Delachaux et Niestle S.A. 3., 1999. — 70 p.
• Kerney M.P., Cameron R.A., Jungbluth J.H. The Land Snails of North and Middle Europe. Hamburg, 1983. 384 p.
• Likharev I.M., Rammel’meyyer E.S. Nazemnyye mollyuski fauny SSSR. M., L.: Izd-vo AN SSSR, 1952. 512 p. (Opredeliteli po faune SSSR. Tr. ZIN AN SSSR. T. 43) (in Russian).
• Lisiecki, L.E., Raymo, M. A Pliocene-Pleistocene stack of 57 globally distributed benthic D18O records// Paleoceanography. 2005. V. 20. P. 1–17. DOI.org / 10.1029 / 2004PA001071
• Lozek V. Quartärmollusken der Tschechoslowakei. Praha, 1964. 374 p. (Rozpravy Ústředního ústavu geologického, Iss. 31).
• Nederlandse Fauna 2. De Nederlandse Zoetwatermollusken. Recente en fossiele weekdieren uit zoet en brak water / Gittenberger, R.E., Janssen, A.W. (Eds.), Nationaal Natuurhistorisch museum Naturalis KNNV uitgeverij. European Invertebrate survey, 1998. 288 p.
• Osipova E., Danukalova G., Marković S. Malacological characteristic of the Middle Pleistocene loess (MIS-6) of the Batajnica locality (Serbia) // Quaternary International. 2013. V. 292. P. 86–100. DOI:10.1016 / j.quaint.2012.10.042
• Osipova E., Danukalova G., Brancaleoni G., Krajcarz T.M., Abdykanova A., Shnaider S. Palaeoenvironmental conditions of the Palaeolithic-Neolithic transition in the Fergana Valley (Central Asia) — New data inferred from fossil molluscs in Obishir-V rockshelter (Kyrgyzstan) // Quaternary International. 2021. V. 605–606. P. 287–299. DOI:10.1016 / j.quaint.2020.11.009
• Puissegur J.‑J. Mollusques continentaux quaternaires de Bourgogne. Significations stratigraphiques et climatiques. Rapports avec d’autres faunes boreales de France // Memoires geologiques de l’Universite de Dijon. 1976. V. 3. 241 p.
• Shileyko A.A. Nazemnyye mollyuski nadsemeystva Helicoidea. L.: Nauka, 1978. 384 p. (Fauna SSSR. Mollyuski. T. III, vyp. 6) (in Russian).
• Shileyko A.A. Nazemnyye mollyuski podotryada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). L.: Nauka, 1984. 399 p. (Fauna SSSR (novaya seriya, №130). T. III, vyp. 3) (in Russian).
• Shileyko A.A., Likharev I.M. Nazemnyye mollyuski semeystva yantarok (Succineidae) fauny SSSR // Sbornik trudov zoologicheskogo muzeya MGU. 1986. T. XXIV. P. 198–238 (in Russian).
• Stratigraficheskiy kodeks Rossii. Izdaniye tret’ye, ispravlennoye i dopolnennoye. SPb.: Izdatel’stvo VSEGEI, 2019. 96 p. (in Russian).
• Sümegi P., Krolopp E. Quartermalacological analyses for modeling of the Upper Weichselian palaeoenvironmental changes in the Carpathian Basin // Quaternary International. 2002. V. 91. P. 53–63. DOI:10.1016 / S1040–6182 (01) 00102–1
• Sysoev A., Shileyko A. Land Snails and Slugs of Russia and Adjacent Countries. Sofia — Moscow: Pensoft Publishers, 2009. 312 p.
• Willis K., Rudher E., Sümegi P. The Full-glacial forests of Central and Southeastern Europe // Quaternary Research. 2000. V. 53. P. 203–213. DOI:10.1006 / qres.1999.2119
• Yakovlev A.G., Danukalova G.A., Alimbekova L.I., Satayev R.M., Nurmukhametov I.M., Makarova O.V. Biostratigraficheskaya kharakteristika geologicheskogo pamyatnika prirody «Peshchera Nukatskaya» // Pleystotsenovyye i golotsenovyye fauny Urala. Chelyabinsk: Rifey Press, 2000. P. 81–104 (in Russian).
• Yakovlev A.G., Danukalova G.A., Alimbekova L.I., Kosintsev P.V., Morozova E.M., Eremeyev A.A. Biostratigraficheskaya kharakteristika golotsenovykh otlozheniy mestonakhozhdeniya Grot Arkheologov (Yuzhnyy Ural) // Geologicheskiy sbornik №3. Informatsionnyye materialy / IG UNTs RAN. Ufa. — 2003. P. 92–98 (in Russian).
• Yakovlev A.G., Danukalova G.A., Yakovleva T.I., Alimbekova L.I., Morozova E.M. Biostratigraficheskaya kharakteristika golotsenovykh otlozheniy mestonakhozhdeniya «Grot Tashmurun» (Yuzhnyy Ural) // Geologicheskiy sbornik №4. — Informatsionnyye materialy / IG UNTs RAN. Ufa. 2004. P. 101–105 (in Russian).
• Yakovlev A.G., Danukalova G.A., Alimbekova L.I., Yakovleva T.I., Eremeyev A.A., Morozova E.M. Biostratigraficheskaya kharakteristika otlozheniy pozdnego neopleystotsena — golotsena v rayone pamyatnika prirody «Vodopad Atysh» (Yuzhnyy Ural) // Fauny Urala i Sibiri v pleystotsene i golotsene. Sb. nauchnykh trudov. Chelyabinsk: Rifey Press, 2005. P. 260–304 (in Russian).
• Yakovlev A., Danukalova G., Kosintcev P., Alimbekova L., Morozova E. Biostratigraphy of the late Palaeolithic site of “Bajslan-tash Cave” (the Southern Urals) // Quaternary International. 2006. V. 149 (1). P. 115–121. DOI:10.1016 / j.quaint.2005.11.025
• Zhadin V.I. Mollyuski presnykh vod SSSR. M.; L.: AN SSSR Press, 1952. 376 p. (Opredeliteli po faune SSSR. Tr. ZIN AN SSSR. T. 46) (in Russian).
• Zilch A., Jaeckel S.G.A. Die Weichtiere (Mollusca) Mitteleuropas / Die Tierwelt Mitteleuropas, Bd. II. L. Iverlag von Quelle and Meyer. Leipzig, 1962. 294 p.